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甘肅鼢鼠(Myospalax cansu)耳蝸結(jié)構(gòu)研究

發(fā)布時(shí)間:2018-06-04 22:01

  本文選題:甘肅鼢鼠 + 耳蝸; 參考:《陜西師范大學(xué)》2010年碩士論文


【摘要】: 地下鼠是由環(huán)境特化的物種,在長(zhǎng)期的地下適應(yīng)過程中,其聽覺系統(tǒng)出現(xiàn)一系列的適應(yīng)性進(jìn)化,其聽覺敏感頻率偏于中低頻,與聲波在地下洞道中的傳播能力相一致。聽覺機(jī)制的核心問題是耳蝸活動(dòng)機(jī)制的研究。 本文運(yùn)用組織學(xué)方法,對(duì)甘肅鼢鼠耳蝸顯微及亞顯微組織結(jié)構(gòu)作了研究,結(jié)果表明: 甘肅鼢鼠耳蝸有3.5圈螺旋形構(gòu)造,這一點(diǎn)和其它哺乳類動(dòng)物相似。從蝸底到蝸頂,均可清楚地看見螺旋神經(jīng)節(jié),螺旋神經(jīng)節(jié)分為大、小兩種類型,數(shù)目從蝸底到蝸頂呈逐漸增多趨勢(shì)。 甘肅鼢鼠耳蝸各圈基底膜上內(nèi)毛細(xì)胞簡(jiǎn)單線形排列為l排,約40根。甘肅鼢鼠耳蝸蝸底基底膜上,OHC為3排,其上聽纖毛呈弧形排列為4排,每排大約30根,每束大約120根左右,聽纖毛由內(nèi)向外逐漸增高,排列整齊,形態(tài)完整。IHC聽纖毛較OHC聽纖毛長(zhǎng),其OHC最內(nèi)排聽纖毛長(zhǎng)度大約1μm,最外排聽纖毛長(zhǎng)度大約為4μm;IHC聽纖毛長(zhǎng)約7μm。OHC之間間隙很小,各排OHC之間鑲嵌排列,各細(xì)胞之間幾乎粘連。外毛細(xì)胞靜纖毛的長(zhǎng)度靠近耳蝸軸最短,靠近耳蝸側(cè)壁最長(zhǎng),由蝸底到蝸頂逐漸變長(zhǎng),靜纖毛的數(shù)目由蝸底到頂逐漸減少。耳蝸蝸頂外毛細(xì)胞和靜纖毛常排列不規(guī)則,而且可見少數(shù)靜纖毛自然缺失。內(nèi)毛細(xì)胞和靜纖毛一直排列到耳蝸頂盲端,毛細(xì)胞和靜纖毛數(shù)目均未減少。從而證明內(nèi)毛細(xì)胞感知低頻信號(hào),長(zhǎng)期地下生活使耳蝸發(fā)生適應(yīng)性進(jìn)化。
[Abstract]:The underground mouse is a special species of environment. During the long period of underground adaptation, a series of adaptive evolution has occurred in its auditory system. The frequency of auditory sensitivity is more moderate and low frequency, which is consistent with the propagation ability of sound wave in underground tunnel. The core problem of auditory mechanism is the study of cochlear mechanism. The microstructures and submicroscopic structures of the cochlea of Gansu zokor were studied by histological method. The results showed that: Gansu zokor cochlea has a 3.5-circle spiral structure, which is similar to other mammals. The spiral ganglion can be clearly seen from the base of the cochlea to the apex of the cochlea. The spiral ganglion can be divided into two types, the number of which is increasing gradually from the cochlea base to the cochlea apex. The hair cells in the basal membrane of the cochlea of Gansu zokor were arranged in a simple line, about 40. The OHC of the basal membrane of cochlea of Gansu zokor is 3 rows. The upper auditory cilia are arranged in 4 rows, about 30 in each row, about 120 in each bundle. The auditory ciliates gradually increase from inside to outside and are arranged neatly. The length of auditory cilium of OHC was about 1 渭 m, and the length of auditory cilium was about 4 渭 m. The gap between auditory cilia was about 7 渭 m.OHC, and the arrangement of OHC was inlaid and almost adherent between cells. The length of the outer hair cells was the shortest near the cochlear axis, the longest near the lateral wall of the cochlea, the longer from the cochlear base to the cochlea apex, and the number of the static cilia gradually decreased from the cochlear base to the top. The outer hair cells and static cilia of cochlea are often arranged irregularly, and a few of them are absent naturally. The number of hair cells and static ciliates was not decreased when the inner hair cells and static cilia were arranged all the way up to the top of the cochlea. It is proved that the inner hair cells perceive low frequency signals and long-term underground life makes cochlea adaptive evolution.
【學(xué)位授予單位】:陜西師范大學(xué)
【學(xué)位級(jí)別】:碩士
【學(xué)位授予年份】:2010
【分類號(hào)】:R764

【參考文獻(xiàn)】

相關(guān)期刊論文 前10條

1 楊軍;脊椎動(dòng)物毛細(xì)胞的機(jī)械換能作用——靜纖毛束的結(jié)構(gòu)與功能[J];國(guó)外醫(yī)學(xué).耳鼻咽喉科學(xué)分冊(cè);1996年06期

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